"We regret the death of Raul Isaias Baduel of cardiorespiratory arrest as a consequence of Covid-19," prosecutor general Tarek Saab posted on Twitter.

Baduel, 66, had received an initial Covid-19 vaccination and "was receiving appropriate medical care," Saab added.

Baduel was Chavez's defense minister, and helped reinstate Chavez after the April 2002 coup that briefly removed the president from power.

The minister served almost eight years on corruption charges. After his release in 2015, he was jailed again on charges of conspiracy against current President Nicolas Maduro.

Baduel's death means "there are now 10 political prisoners who have died in custody" since 2015, said Gonzalo Himiob, a lawyer for the human rights NGO Foro Penal.

"The responsibility for the life and health of any detainee rests with the state. Medical treatment is continuously required for prisoners," Himiob said on Twitter.

Two of Baduel's sons have also been arrested on allegations of conspiracy. One, Raul Emilio, is at large.

The other, Josnars Adolfo Baduel, was arrested for alleged participation in a failed maritime invasion in 2019 that sought to overthrow Maduro. He is still in custody.

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